Millipedes of Australia

Sphaerotheriida: Epicyliosoma (Paracyliosoma) spp.

Go to Epicyliosoma synonymy on Epicyliosoma introduction page

Epicyliosoma (Paracyliosoma) denticulatum (Verhoeff, 1924)
  Cyliosoma denticulatum Verhoeff 1924: 45 (in species key),
    47 (first description),
    figs. 23, 24, 27, 28, pl. 2.
  Cyliosoma denticulatum Verhoeff 1928: 81. (In species keys.)
  Cyliosoma denticulatum Jeekel 1981: 26. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) denticulatum Jeekel 1986: 37 (new combination),
    38 (in species key).

Syntypes. 2 males and 1 juvenile female.
Locality. Blackall Range, Qld.
Collector, date. Collected during Dr E. Mjöberg's Australia expeditions, 1910-1913.
Deposited. NHRM?

Epicyliosoma (Paracyliosoma) mjoebergi (Verhoeff, 1924)
  Cyliosoma queenslandicum mjöbergi Verhoeff 1924: 45 (in species key as subspecies of 'C. queenslandicum', which is Verhoeff's renaming of C. queenslandiae Brolemann, 1913),
    47 (first description, as subspecies of 'C. queenslandicum').
  Cyliosoma queenslandicum mjöbergi Verhoeff 1928: 80, 81. (In species keys.)
  Cyliosoma queenslandiae mjoebergi Jeekel 1981: 24. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) mjoebergi Jeekel 1986: 37 (raised to species in new combination),
    38 (in species key).

Syntypes. 7 specimens, among them 1 male from Colosseum, Qld, November (Verhoeff 1924: 47).
Collector, date. Collected during Dr E. Mjöberg's Australia expeditions, 1910-1913.
Deposited. No information yet. NHRM?
Note. 'E. mjoebergi was described briefly as a subspecies of E. queenslandicum. However, the relationship of mjoebergi to queenslandicum is dubious, because the latter was probably based on material of more than one species. Rather mjoebergi seems related to, or perhaps even identical with E. queenslandiae.' (Jeekel 1986: 37)

Epicyliosoma (Paracyliosoma) penicilligerum (Verhoeff, 1928)
  Cyliosoma (Paracyliosoma) penicilligerum Verhoeff 1928: 80, 81 (in species keys),
    83 (first description),
    84 (in species key),
    figs. 5-7, pl. 6.
  Cyliosoma penicilligerum Jeekel 1971: 27. (In genus catalogue as type of Paracyliosoma.)
  Cyliosoma penicilligerum Jeekel 1981: 26. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) penicilligerum Jeekel 1986: 36 (in species list as Cyliosoma penicilligerum, type of Paracyliosoma),
    37 (new combination),
    38 (in species key).

Syntypes. 'A female and four males from North Dorrigo [NSW], collected by A. Musgrave' (Verhoeff 1928: 83).
Deposited. AM, KZ 76522; this is a female collected from North Dorrigo on 4 January 1923 by A. Musgrave and labelled 'lectopapatype female', probably by P.M. Johns (G. Milledge, AM, pers. comm.).

Epicyliosoma (Paracyliosoma) queenslandiae (Brolemann, 1913)
  Cyliosoma queenslandiae Brolemann 1913: 80 (first description),
    figs. 1-4, pl. 14; fig. 14, p. 78; fig. 15, p. 83; figs. 16-21, p. 84.
  Cyliosoma Queenslandiae Attems 1914: 142. (In species catalogue.)
  Cyliosoma Queenslandiae Silvestri 1917: 62 (in species list, but Silvestri adds '=Cyliosoma unicolor Silv.'),
    71 (synonymised with Cyliosoma unicolor).
  Cyliosoma queenslandiae Chamberlin 1920a: 87. (In species catalogue as synonym of Cyliosoma unicolor.)
  Cyliosoma queenslandiae Verhoeff 1924: 45. (Redescribed as 'Cyliosoma queenslandicum Bröl. (= queenslandiae Bröl.)' from new material.)
  Cyliosoma queenslandiae Jeekel 1981: 24. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) queenslandiae Jeekel 1986: 37 (new combination),
    38 (in species key).

Syntypes. 4 males and 3 females (Brolemann 1913: 83).
Locality. Gayndah, Qld.
Collector, date. No information yet.
Deposited. AM holds 4 samples (G. Milledge, AM, pers. comm.):
  KS 76517, labelled 'lectotype male, P.M. Johns 14 XII 67'
  KS 76518, labelled 'lectoparatypes, 3 males, P.M. Johns 14 XII 67'
  KS 76519, labelled 'lectoparatypes, 3 females, P.M. Johns 14 XII 67'
  KS 76520, no label, unclear what these specimens are.
Johns has not published this lectotypification, which is therefore not valid. The discrepancy in number of specimens is unexplained.
Note. 'E. queenslandiae was synonymized by Silvestri (1917) with E. unicolor. Indeed there seems to be an almost complete agreement in the characters of the two, and the fact that they were described from the same locality further supports synonymy. The very long prefemoral process of the anterior telopods described for unicolor is sufficient reason to keep the two separate for the time being.' (Jeekel 1986: 37)

Epicyliosoma (Paracyliosoma) queenslandicum (Verhoeff, 1924)
  Cyliosoma queenslandicum Verhoeff 1924: 45 (first description as 'Cyliosoma queenslandicum Bröl. (= queenslandiae Bröl.)' from new material),
    figs. 29, 30, pl. 2.
  Cyliosoma queenslandicum Verhoeff 1928: 80, 81 (in species keys),
    84 (redescription, species key).
  Epicyliosoma (Paracyliosoma) queenslandicum Jeekel 1986: 37 (new combination),
    38 (in species key as 'targionii queenslandicum').

Syntypes. A dozen specimens of both sexes ('Untersucht wurden ein Dutzend Individuen beider Geschlechter'; Verhoeff 1924: 47).
Localities. Herberton, Mt Tamborine and Malanda, Qld.
Collector, date. Collected during Dr E. Mjöberg's Australia expeditions, 1910-1913.
Deposited. No information yet. NHRM?
Note. 'E. queenslandicum may be identical with E. targionii. It seems likely, however, that Verhoeff mixed up two species under the name: one from northern Queensland and one from southern Queensland, subsequently reported also from northern New South Wales. A restudy of the type material, involving also the selection of a lectotype, is needed to clarify this point. It must be noted here, that the name queenslandicum is in fact a replacement name for queenslandiae and as such an objective synonym of the latter. It is doubtful that the material studied by Brölemann and Verhoeff belongs to one species. On that account the name queenslandicum is maintained here as a matter of convenience and in order not to complicate matters.' (Jeekel 1986: 37)

Epicyliosoma (Paracyliosoma) targionii (Silvestri, 1898)
  Cyliosoma Targionii Silvestri 1898: 226 (first description),
    figs. 1-3, p. 226.
  Cyliosoma targionii Brolemann 1913: 80 (in species list),
    83 (compares with C. queenslandiae),
    84 (reproduces Silvestri's first description).
  Cyliosoma Targionii Attems 1914: 142. (In species catalogue.)
  Cyliosoma (Cyliosoma) Targionii Silvestri 1917: 62 (in species list),
    69 (redescription),
    fig. 1, p. 63; figs. 2, 3, p. 64; fig. 4, p. 65; fig. 5, p. 66.
  Cyliosoma targioni Chamberlin 1920a: 87. (In species catalogue.)
  Cyliosoma targionii Jeekel 1981: 24. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) targionii Jeekel 1986: 37 (new combination),
    38 (in species key as 'targionii queenslandicum').

Types. Male and female syntypes.
Locality. Cairns, Qld.
Collector, date. [Giovanni] Podenzana, December 1891 (Silvestri 1898: 225).
Deposited. MSNF (Silvestri 1898: title).
Note. 'E. queenslandicum may be identical with E. targionii. It seems likely, however, that Verhoeff mixed up two species under the name: one from northern Queensland and one from southern Queensland, subsequently reported also from northern New South Wales. A restudy of the type material, involving also the selection of a lectotype, is needed to clarify this point. It must be noted here, that the name queenslandicum is in fact a replacement name for queenslandiae and as such an objective synonym of the latter. It is doubtful that the material studied by Brölemann and Verhoeff belongs to one species. On that account the name queenslandicum is maintained here as a matter of convenience and in order not to complicate matters.' (Jeekel 1986: 37)

Epicyliosoma (Paracyliosoma) unicolor (Silvestri, 1897)
  Cyliosoma unicolor Silvestri 1897a: 16 (first description),
    figs. 80-82, pl. 2.
  Cyliosoma unicolor Brolemann 1913: 80 (in species list),
    84 (compares with C. queenslandiae, reproduces Silvestri's first description).
  Cyliosoma unicolor Attems 1914: 142. (In species catalogue.)
  Cyliosoma (Cyliosoma) unicolor Silvestri 1917: 62 (in species list),
    68 (in species key),
    71 (redescription),
    fig. 7, p. 71.
  Cyliosoma unicolor Chamberlin 1920a: 87. (In species catalogue.)
  Cyliosoma unicolor Jeekel 1981: 24. (In list of species recorded from eastern Australia.)
  Epicyliosoma (Paracyliosoma) unicolor Jeekel 1986: 37 (new combination),
    38 (in species key).

Syntypes. At least 1 male and 1 female.
Locality. Gayndah, Qld.
Collector, date. No information yet.
Deposited. SNSD; no. 283 (Silvestri 1897a: 16).
Note. 'E. queenslandiae was synonymized by Silvestri (1917) with E. unicolor. Indeed there seems to be an almost complete agreement in the characters of the two, and the fact that they were described from the same locality further supports synonymy. The very long prefemoral process of the anterior telopods described for unicolor is sufficient reason to keep the two separate for the time being.' (Jeekel 1986: 37)