Tasmanian Millipedes

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Polydesmida
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Order Polydesmida: Identification (part 1)

It is hard to write an identification key for Polydesmida which is not based largely on their gonopods. Below is my first attempt at such a key, and it may not work as smoothly as users would like. If you find yourself getting lost and frequently backtracking, let me know the problems by email, and I will try to improve the key.

The key mainly requires you to look carefully at the paranota (singular paranotum), which are lateral extensions of the tergites (p in image at right).
 
You should also look carefully at the tiny spiracles, through which a millipede breathes. On most segments there are two spiracles on each side, opening just above the leg bases (s in image at right).
 
Finally, pay attention to the ozopores. These are openings to the chemical defence glands. In Polydesmida they open on the side of the body, one per segment, and usually on segments 5, 7, 9, 10, 12, 13, and 15-18 (H+19) or 15-19 (H+20). This is called the 'normal pore formula'. The size, shape, position and number of the ozopores can vary from genus to genus.
 
Note: For cave-dwelling H+20 millipedes from Ida Bay and Precipitous Bluff, go straight to the Atalopharetra page.

paranota
spiracles

 

1a. Paranota broad and obvious;
                go to 2

broad paranota

1b. Paranota greatly reduced or absent;
                go to 10 in part 2 of the key

no paranota
 

2a. Rear corner of paranotum clearly projecting backwards (left image);
                go to 3
 
2b. Rear corner of paranotum square or slightly rounded (right image);
                go to 6

projectionno projection
 

3a. Prominent long seta arising near rear corner of paranotum;
                go to 4
 
3b. No such seta;
                go to 5

PCS
 

4a. Spiracles large, seemingly swollen (except Dasystigma tyleri, q.v.) adult 20-25 mm long, pale brown or chestnut brown;
                Dasystigma spp.
 
4b. Spiracles large, seemingly swollen, close together; adult >30 mm long, red-brown or purplish-brown, NE only;
                Tasmanopeltis grandis

Dasystigma
 

4c. Spiracles inconspicuous, not seemingly swollen, not close together;
                Lissodesmus spp.

Lissodesmus
 

5a. Adults at least 15 mm long, purplish-brown; tergites flat; found in bush areas;
                Tasmanodesmus hardyi
 
5b. Adults no more than 10 mm long, pale brown; tergites with large, parallel swellings; mainly gardens;
                Brachydesmus superus

 

6a. Second tergite greatly expanded laterally (see image at right), tergites covered with numerous tiny bumps, adults 5-6 mm long;
                Asphalidesmus spp.
 
6b. Second tergite not greatly expanded, tergites not covered with numerous tiny bumps;
                go to 7

Asphalidesmus
 

7a. Tergites in adults smooth, sometimes with 3 tranverse rows of low, round swellings; adults generally with reddish pigmentation;
                Tasmaniosoma and the ER group
 
7b. Tergites in adults with 3 tranverse rows of small setae; sometimes coloured but never reddish;
                go to 8

 

8a. Body usually well-pigmented (various colours); males with gonopods with large, rearwards-projecting 'flange' (see image at right);
                Atrophotergum spp.
 
8b. Body white or faintly pigmented; mainly tiny species 4-6 mm long as adults;
                Go to 9

Atrophotergum
 

9a. Males with long gonopods jointed in middle (see image at right);
                Ginglymodesmus spp.
 
9b. Gonopods without joint in middle
                F, S and T miscellany;

Ginglymodesmus

Click here to go to key couplet 10.